O A MISUND
Institute of Marine Research, P.O. Box 1870, N-5817 Bergen, Norway
J C COETZEE
Marine & Coastal Management, Department of Environmental Affairs and Tourism, Private Bag X2, Rogge Bay 8012, Cape Town, South Africa
P FRÉON
Institut de Recherche pour le Dèveloppement, France, and Marine & Coastal Management, Cape Town, South Africa
M GARDENER
Institute of Maritime Technology, Simonstown, South Africa
K OLSEN
Norwegian College of Fishery Science, University of Tromsø, N-9037 Tromsø, Norway
I SVELLINGEN
Institute of Marine Research, P.O. Box 1870, N-5817 Bergen, Norway
I HAMPTON
Fisheries Resource Surveys, 30 Jeffcoat Avenue, Bergvliet 7945, South Africa
Abstract
The schooling behaviour of sardine Sardinops sagax in False Bay on the south coast of South Africa was studied in October 1995 using high-resolution sonar and a hull-mounted, echo-integration unit. School formation and disintegration were studied, and school shape, packing density, swimming behaviour and inter- and intra-school event rates were quantified. Mean fish density was 29.5 fish m-3 (SD 46 fish m-3), but it varied between schools by a factor of about 100 (from 2 to 233 fish m-3). Tracked schools moved at average speeds of 0.67–1.59 m s-3. Schools changed shape on average every 2.08 minutes, and underwent either splits or merges with other schools on average every 5 minutes. Relationships between the geometric dimensions and biomass of the schools were established.
Afr. J. mar. Sci. 25: 185–193